Animals
What is an Animal?
Animals are:
Eukaryotic
Multicellular
Heterotrophic organisms that obtain nutrients by ingestion
Able to digest their food within their bodies
Animal cells lack the cell walls that provide strong support in the bodies of plants and fungi.
Most animals:
Have muscle cells
Have nerve cells that control the muscles
Are diploid
Reproduce sexually
Proceed through a series of typically similar developmental stages
Early Animals and the
Cambrian Explosion
Animals probably evolved from a colonial flagellated protist that lived in Precambrian seas about 600–700 million years ago.
At the beginning of the Cambrian period, 542 million years ago, animals underwent a rapid diversification.
During a span of about 15 million years:
All major animal body plans we see today evolved
Many of these animals seem bizarre
Animal Phylogeny
Biologists categorize animals by:
General features of body structure
More recently, using genetic data
One major branch point distinguishes sponges from all other animals because, unlike more complex animals, sponges lack true tissues.
A second major evolutionary split is based on body symmetry.
Radial symmetry refers to animals that are identical all around a central axis.
Bilateral symmetry exists where there is only one way to split the animal into equal halves.
Animals also vary according to the presence and type of body cavity, a fluid-filled space separating the digestive tract from the outer body wall.
There are differences in how the body cavity forms.
If the body cavity is not completely lined by tissue derived from mesoderm, it is a pseudocoelom.
A true coelom is completely lined by tissue derived from mesoderm.
Sponges
Sponges represent multiple phyla.
Sponges include sessile animals that lack true tissues and that were once believed to be plants.
The body of a sponge resembles a sac perforated with holes.
Choanocyte cells draw water through the walls of the sponge where food is collected.
Cnidarians
Cnidarians (phylum Cnidaria) are characterized by:
The presence of body tissues
Radial symmetry
Tentacles with stinging cells
The basic body plan of a cnidarian is a sac with a gastrovascular cavity, a central digestive compartment with only one opening.
The body plan has two variations:
The sessile polyp
The floating medusa
Cnidarians are carnivores that use tentacles, armed with cnidocytes (“stinging cells”), to capture prey.
Molluscs
Molluscs (phylum Mollusca) are represented by soft-bodied animals, usually protected by a hard shell.
Many molluscs feed by using a file-like organ called a radula to scrape up food.
The body of a mollusc has three main parts:
A muscular foot used for movement
A visceral mass housing most of the internal organs
A mantle, which secretes the shell if present
The three major groups of molluscs are:
Gastropods, protected by a single, spiraled shell.
Bivalves, with a shell divided into two halves hinged together.
Cephalopods, which typically lack an external shell and are built for speed and agility.
Flatworms
Flatworms (phylum Platyhelminthes) are the simplest bilateral animals.
Flatworms include forms that are:
Parasites or
Free-living in marine, freshwater, or damp habitats
The gastrovascular cavity of flatworms is highly branched
Provides an extensive surface area for absorption of nutrients
Annelids
Annelids (phylum Annelida) have:
Body segmentation, a subdivision of the body along its length into a series of repeated parts
A coelom
A complete digestive tract with two openings (a mouth and anus), and one-way movement of food.
The three main groups of annelids are:
Earthworms, which eat their way through soil
Polychaetes, marine worms with segmental appendages for movement and gas exchange
Leeches, typically free-living carnivores but with some bloodsucking forms
Roundworms
Roundworms (phylum Nematoda) are:
Cylindrical in shape, tapered at both ends
The most diverse and widespread of all animals
Roundworms (also called nematodes) are:
Important decomposers
Dangerous parasites in plants, humans, and other animals
Arthropods
Arthropods (phylum Arthropoda) are named for their jointed appendages.
There are about one million arthropod species identified, mostly insects.
Arthropods are a very diverse and successful group, occurring in nearly all habitats in the biosphere.
There are four main groups of arthropods.
General Characteristics of Arthropods
Arthropods are segmented animals with specialized segments and appendages for an efficient division of labor among body regions.
The body of arthropods is completely covered by an exoskeleton, an external skeleton that provides:
Protection
Points of attachment for the muscles that move appendages
Arachnids:
Live on land
Usually have four pairs of walking legs and a specialized pair of feeding appendages
Include spiders, scorpions, ticks, and mites
Crustaceans:
Are nearly all aquatic
Have multiple pairs of specialized appendages
Include crabs, lobsters, crayfish, shrimps, and barnacles
Millipedes:
Eat decaying plant matter
Have two pairs of short legs per body segment
Centipedes:
Are terrestrial carnivores with poison claws
Have one pair of short legs per body segment
Insects typically have a three-part body:
Head
Thorax
Abdomen
The insect head usually bears:
A pair of sensory antennae
A pair of eyes
The mouthparts are adapted for particular kinds of eating.
Flight is one key to the great success of insects.
Insects outnumber all other forms of life combined.
Insects live in:
Almost every terrestrial habitat
Freshwater
The air
Many insects undergo metamorphosis in their development.
Young insects may:
Appear to be smaller forms of the adult or change from a larval form to something much different as an adult
Echinoderms
Echinoderms (phylum Echinodermata):
Lack body segments
Typically show radial symmetry as adults but bilateral symmetry as larvae
Have an endoskeleton
Have a water vascular system that facilitates movement and gas exchange
Echinoderms are a very diverse group.
Characteristics of
Chordates
Chordates (phylum Chordata) all share four key features that appear in the embryo and sometimes the adult:
A dorsal, hollow nerve cord
A notochord
Pharyngeal slits
A post-anal tail
Another chordate characteristic is body segmentation, apparent in the:
Backbone of vertebrates
Segmental muscles of all chordates
Chordates consists of three groups of invertebrates:
Lancelets are bladelike animals without a cranium.
Tunicates, or sea squirts, also lack a cranium.
Hagfishes are eel-like forms that have a cranium.
All other chordates are vertebrates.
Fishes
The first vertebrates were aquatic and probably evolved during the early Cambrian period, about 542 million years ago. They:
Lacked jaws
Are represented today by hagfishes
Lampreys:
Are vertebrates
Have a cranium
But lack jaws
The two major groups of living fishes are the:
Cartilaginous fishes (sharks and rays) with a flexible skeleton made of cartilage
And bony fishes with a skeleton reinforced by hard calcium salts
Bony fishes include:
Ray-finned fishes
Lungfishes
Lobe-finned fishes
Cartilaginous and bony fishes have a lateral line system that detects minor vibrations in the water.
To provide lift off the bottom:
Cartilaginous fish must swim but
Bony fish have swim bladders, gas-filled sacs that make them buoyant
Amphibians
Amphibians:
Exhibit a mixture of aquatic and terrestrial adaptations
Usually need water to reproduce
Typically undergo metamorphosis from an aquatic larva to a terrestrial adult
Amphibians:
Were the first vertebrates to colonize land
Descended from fishes that had lungs and fins with muscles
Terrestrial vertebrates are collectively called tetrapods, which means “four feet.”
Tetrapods include:
Amphibians
Reptiles
Mammals
Reptiles
Reptiles (including birds) and mammals are amniotes, which produce amniotic eggs that consist of a fluid-filled shell inside of which the embryo develops.
Reptiles include:
Snakes
Lizards
Turtles
Crocodiles
Alligators
Birds
Reptile adaptations to living on land include:
Amniotic eggs
Scaled, waterproof skin
Non-bird reptiles are ectotherms, sometimes referred to as “cold-blooded,” which means that they obtain their body heat from the environment.
A non-bird reptile can survive on less than 10% of the calories required by a bird or mammal of equivalent size.
Reptiles diversified extensively during the Mesozoic era.
Dinosaurs were the largest animals ever to live on land.
Recent genetic evidence shows that birds evolved from a lineage of small, two-legged dinosaurs during the great reptilian radiation of the Mesozoic era.
Birds have many adaptations that make them lighter in flight:
Honeycombed bones
One instead of two ovaries
A beak instead of teeth
Unlike other reptiles, birds are endotherms, maintaining a warmer and steady body temperature.
Birds wings adapted for flight are airfoils, powered by breast muscles anchored to a keel-like breastbone.
Mammals
The first true mammals:
Arose about 200 million years ago
Were probably small, nocturnal insect-eaters
Most mammals are terrestrial although dolphins, porpoises, and whales are totally aquatic.
Mammalian hallmarks are:
Hair
Mammary glands that produce milk, which nourishes the young
There are three major groups of mammals:
Monotremes, egg-laying mammals
Marsupials, pouched mammals with a placenta
And eutherians, placental mammals
Eutherian placentas provide more intimate and long-lasting association between the mother and her developing young than do marsupial placentas.
The Evolution of
Primates
Primates evolved from insect-eating mammals during the late Cretaceous period.
Primates are distinguished by characteristics that were shaped by the demands of living in trees. These characteristics include:
Limber shoulder joints
Eyes in front of the face
Excellent eye-hand coordination
Extensive parental care
Taxonomists divide the primates into three main groups.
The first group of primates includes:
Lorises
Pottos
Lemurs
Tarsiers form the second group.
The third group, anthropoids, includes monkeys.
Hominoids consist of humans and their ape relatives.
The Emergence of Humankind
Humans and chimpanzees have shared a common African ancestry for all but the last 5–7 million years.
Some Common
Misconceptions
Chimpanzees and humans represent two divergent branches of the anthropoid tree that each evolved from a common, less specialized ancestor.
Our ancestors were not chimpanzees or any other modern apes.
Human evolution is not a ladder with a parade of fossil hominids (members of the human family) leading directly to modern humans.
Instead, human evolution is more like a multi-branched bush than a ladder.
At times in hominid history, several different human species coexisted.